passages.id: split_1_test_1347_passage
passages.type: progene_text
passages.text: By matrix - assisted laser desorption ionization - time of flight mass spectrometry , two alkaline phosphatases ( APases ) ( PhoA and PhoB ) , an APase - alkaline phosphodiesterase ( PhoD ) , a glycerophosphoryl diester phosphodiesterase ( GlpQ ) , and the lipoprotein YdhF were identified as very strongly induced PhoPR - dependent proteins secreted into the extracellular medium .